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Biomedical subjects

D M Weary

Publications and source records attributed to D M Weary.

At least 19 recordsLinked to original sources

Differential effects of sodium and magnesium sulfate on water consumption by beef cattle.

The existing guidelines for maximum sulfate (SO4) in cattle drinking water are based on Na2SO4, although many water sources contain greater concentrations of MgSO4. Two experiments compared the effect of different SO4 salts on water consumption and fecal DM of cattle. In Exp. 1, 8 yearling heifers (initial BW = 345 +/- 8 kg; mean +/- SD) were watered twice daily with tapwater or water containing Na2SO4 or MgSO4 at target levels of 1,500, 3,000, or 4,500 mg of SO4/L for 2-d treatment periods separated by 2 d of access to tapwater. In Exp. 2, 16 yearling cattle (initial BW = 421 +/- 24 kg) were watered twice daily with tap-water (16 mg of SO4/L) or water containing Na2SO4 at target levels of 2,000 mg of SO4/L (low Na2SO4), MgSO4 at 2,000 mg of SO4/L (low MgSO4), or MgSO4 at 4,000 mg of SO4/L (high MgSO4) in 21-d treatment periods separated by 7-d periods on tapwater. The first 10 d of each period were allowed for adjustment to the treatment, and the final 11 d was considered the treatment period for analysis purposes. Treatments were applied in an incomplete Latin square, where each animal was exposed to 3 of the 4 treatments. In Exp. 1, the average daily water consumption decreased linearly as the SO4 concentration increased for MgSO4 (P = 0.0001) but not for Na2SO4 (P = 0.39). In Exp. 2, the average daily water consumption was less for cattle on the high-MgSO4 treatment than for cattle on the low-MgSO4 treatment (P = 0.0001), and cattle on the low-MgSO4 treatment tended (P = 0.09) to drink less than those on the tap-water treatment. Fecal DM was greater for cattle on the high-MgSO4 treatment than for those on the low-MgSO4 treatment (P < 0.01). These findings indicate that cattle reduce their consumption of water containing high (> or = 4,000 mg of SO4/L) concentrations of MgSO4, even after a given time to adjust to the treatment; such reductions may be accompanied by an increase in fecal DM.

Animals↗

Effect of hoof pathologies on subjective assessments of dairy cow gait.

To explore how hoof pathologies affect dairy cattle gait, we studied cows with sole hemorrhages (n = 14), sole ulcers (n = 7), and those with no visible injuries (n = 17). Overall gait assessments, scored from video using a 1 to 5 numerical rating system (1 = sound, 5 = severely lame) and a continuous 100-unit visual analog scale, found cows having sole ulcers had poorer gait than healthy cows (mean +/- SEM: 4.0 +/- 0.13 vs. 3.1 +/- 0.08, and 59 +/- 3 vs. 46 +/- 2, respectively). Six gait attributes (back arch, head bob, tracking-up, joint flexion, asymmetric gait, and reluctance to bear weight) were also assessed using continuous 100-unit scales. Compared with healthy cows, those having sole ulcers walked with a more pronounced back arch (12 +/- 3 vs. 28 +/- 4), more jerky head movement (2 +/- 2 vs. 10 +/- 3), shortened strides (7 +/- 2 vs. 26 +/- 4), and more uneven weighting among the limbs (16 +/- 2 vs. 32 +/- 3). Of all measures, the numerical rating system most effectively discriminated healthy cows from those with sole ulcers (R2 = 0.73), classifying 92% of animals correctly. No differences were detected among cows with and without sole hemorrhages. Intra- and interobserver reliabilities were reasonable for all measures (R2 > or = 0.64) except joint flexion and asymmetric gait. In summary, subjective assessments of dairy cattle gait provide valid and reliable approaches to identifying cattle with sole ulcers.

Animals↗

Effects of mixing on drinking and competitive behavior of dairy calves.

Group housing provides increased access to space and social interactions for calves while reducing labor costs for producers. However, group housing necessarily requires that calves be mixed and no research to date has addressed the effects of mixing on behavior of milk-fed dairy calves. The objective of this study was to monitor the feeding and competitive behavior of individual dairy calves (n = 8) after introduction into an established group of older calves fed ad libitum by a computer-controlled milk feeder. Milk feeding was monitored for 2 d before introduction into the new group and both milk feeding and competitive behaviors were monitored for 4 d after mixing. Mean (+/- SE) milk consumption before mixing was 9.7 +/- 0.7 kg/d, dropped slightly on the day of mixing to 8.6 +/- 0.6 kg/d, but increased on d 1 to 3 after mixing to 11.1 +/- 0.3 kg/d. Calves visited the feeder less frequently on the day of mixing (6.0 +/- 1.8 visits/d) than on either the days before mixing (20.3 +/- 2.5 visits/d) or the days after mixing (25.3 +/- 6.9 visits/ d). The mean duration of feeder visits and mean milk consumption per visit increased from 4 min 15 s +/- 21 s and 0.53 +/- 0.06 kg per visit before mixing to 8 min 17 s +/- 1 min 28 s and 1.87 +/- 0.51 kg per visit on the day of mixing. Competitive displacements from the milk-feeding stall were rare. In summary, feeding behavior of young calves is altered on the day of mixing, but calves are able to maintain milk intake when using a milk feeder fitted with a stall that prevents calves from displacing one another.

Aggression↗

Flooring in front of the feed bunk affects feeding behavior and use of freestalls by dairy cows.

In 2 experiments we assessed how preferences, time budgets, and feeding behavior of dairy cows change in response to flooring surfaces in front of the feed bunk. In Experiment 1, 12 nonlactating dairy cattle were individually housed with access to 2 standing platforms filled with either concrete or sawdust. In Experiment 2, 24 nonlactating dairy cattle were given access to either concrete or Animat rubber flooring in front of the feed bunk. In Experiment 1, cows preferred the sawdust to the concrete flooring. In both experiments, cows provided with a softer floor in front of the feed bunk spent more time standing near the feed bunk without eating (Experiment 1: 67 vs. 40 min/d on sawdust vs. concrete, respectively, SEM = 5.6 min/d; Experiment 2: 176 vs. 115 min/d on Animat vs. concrete, respectively, SEM = 20.5 min/d) compared with when they were kept on concrete. The increased time spent at the feed bunk was due to a combination of more frequent eating and standing bouts, indicating that cows were more willing to move on nonconcrete flooring. Total time spent eating was significantly greater on the softer floor in Experiment 2, but not in Experiment 1 (Exp. 1: 289 vs. 275 min/d on sawdust and concrete, respectively, SEM = 7.3 min/d; Exp. 2: 330 vs. 289 min/d on Animat and concrete, respectively, SEM = 15.4), although feed intake was increased on the sawdust treatment in Experiment 1. Cows spent significantly more time lying in the feed alley when the flooring was rubber (219 vs. 53 min/d on Animat and concrete, SEM = 53.6 min/d), perhaps because the lying area in Experiment 2 was inadequate. In conclusion, cows prefer to stand on softer flooring in front of the feed bunk, and are more willing to move on and spend more time standing in front of the feed bunk when provided with softer flooring. These results indicate that cows find softer flooring surfaces more comfortable to stand on than concrete, and highlight the importance of evaluating the comfort of the entire facility.

Animal Welfare↗

Effects of milking on dairy cow gait.

We studied cows with (n = 6) and without (n = 26) sole ulcers before and after milking to explore how milking influences dairy cattle gait and how this differs for cows with hoof injuries. Video recordings of cows were digitized using motion-analysis software to calculate stride variables for each hoof. Gait was scored using a 1-to-5 numerical rating system (1 = sound, 5 = severely lame) and a continuous 100-unit visual analog scale of gait attributes (back arch, head bob, tracking-up, and reluctance to bear weight). For cows with and without sole ulcers, differences in gait before and after milking were evident; after milking, all cows had significantly longer strides (123.3 vs. 133.5 +/- 2.0 cm, respectively), higher stride height (8.3 vs. 8.9 +/- 0.1 cm), shorter stride durations (1.49 vs. 1.41 +/- 0.03 s), walked faster (0.85 vs. 0.97 +/- 0.03 m/s), and had shorter periods of triple support (3 legs in ground contact; 80.0 vs. 71.7 +/- 2.0%). Tracking-up and reluctance to bear weight improved after milking (20 vs. 16 +/- 2; 20 vs. 15 +/- 1, respectively), but numerical rating scores and back arch did not. Cows with sole ulcers walked differently than cows without for all measures, except swing duration, both before and after milking. Interactions between hoof health and milking were found for swing duration and head bob but when tested separately, the only effect was that cows without sole ulcers had longer swing durations before milking (0.45 vs. 0.44 +/- 0.01 s, respectively). Gait differences were probably due to udder distention and motivation to return to the home pen. Our results suggest that the most suitable time to assess lameness is after milking when differences between cows with and without ulcers are most evident.

Animals↗

Effects of continuous versus periodic milk availability on behavior and performance of dairy calves.

Dairy calves fed milk ad libitum engage in feeding activity throughout the day and night. However, providing calves with continuous access to milk can present practical problems with milk quality, particularly during warm weather. One way to maintain milk quality in ad libitum feeding regimens is to limit the time that milk is available, but no research to date has addressed how this might affect calves. The objectives of this study were to quantify the effects of continuous access to milk (24 h/d) vs. access during only part of the day on milk feeding behavior and performance. Twenty-eight female Holstein calves were randomly assigned to treatments at 5 +/- 3 d of age and monitored for 28 d. All calves had milk delivered through a teat. Treatments were 1) access to milk for 24 h/d; 2) access to milk for 2 feedings per day (each of 2 h), with water available through the teat the remainder of the day (4-h wet treatment); and, 3) access to milk for 2 feedings per day (each of 2 h), but with no water available through the teat (4-h dry treatment). Calves given 24-h access to milk consumed 11.2 +/- 0.5 kg/d on average. Milk intake tended to be lower for the 4-h wet and 4-h dry treatments, averaging 10.0 +/- 0.3 kg/d, with no difference between these 2 treatments in this or any other variable. Average daily gain over the course of the experiment did not vary with treatment and averaged 1.1 +/- 0.04 kg/d. Calves in the 24-h treatment spent more time on the teat than those calves on the 4-h treatments, but during the 4 h/d when milk was available to all calves, calves in the 4-h treatments spent more time on the teat. These results show that individually housed calves fed milk ad libitum for only 4 h/d compensate by changing their milk feeding behavior and are able to achieve similar weight gains to animals fed milk continuously.

Animal Nutritional Physiological Phenomena↗

Hoof discomfort changes how dairy cattle distribute their body weight.

Lameness is a costly and widespread health and welfare problem in intensive dairy production, and reliable automated methods to detect lameness are needed. Lameness may be detected through the measurement of how cattle distribute their weight among their 4 legs, but this requires an understanding of how cattle redistribute their weight in response to pain in one or more limbs. In 3 experiments, 13, 12, and 15 Holstein dairy cows were trained to stand on a platform that measured the weight placed on each limb. We replaced the soft rubber surface under either 1 or 2 hooves with an uncomfortable concrete surface. Cows placed less weight on a back hoof that was placed on an uncomfortable surface, and they redistributed the majority of the weight onto the contralateral back hoof but did not change the distribution of weight on their front hooves. When the same surface was placed under a front hoof, cows placed less weight on that hoof and placed more weight on the contralateral front hoof and the ipsilateral back hoof. The variation in weight the cow placed on both contralateral hooves increased when one of the hooves was on the uncomfortable surface. Cows placed more weight on the back hooves when both front hooves were standing on uncomfortable surfaces, although no change was observed when back hooves were on uncomfortable surfaces. Dairy cows remove weight from a limb in response to limb discomfort and redistribute this weight primarily to the contralateral limb. The variation in weight over time applied to a pair of contralateral limbs increases in response to discomfort in one hoof. Cows have only limited ability to shift weight from front to back. Measures of weight distribution may provide useful on-farm techniques for the detection of lameness.

Animals↗

Brisket boards reduce freestall use.

We examined how the presence of a brisket board influenced cow preference, stall use, and position within the stall. When given a choice between stalls with or without a brisket board, 15 nonlactating cows spent 68% of their time lying in the stalls without a brisket board, indicating that they preferred this option. When 13 cows had access to either stalls with a brisket board or ones without, they spent, on average, 1.2 h/d more time lying down in stalls without a brisket board. Resting cows positioned themselves relatively forward in the stalls in 98 +/- 5% (mean +/- SE) of lying bouts when the brisket board was absent, compared with 67 +/- 5% of bouts when the board was present. Longer cows were more likely than shorter cows to move forward in the stalls without a brisket board. Cows also had longer lying bouts in stalls without the brisket board (absent: 1.7 +/- 0.08; present: 1.5 +/- 0.08 h/bout). Although it seems likely that the brisket board helps keep stalls clean by positioning cows closer to the curb, our results indicate that brisket boards also make stalls less comfortable for cows. Stall features designed to reduce stall maintenance may compromise cow comfort. We suggest that new approaches to cow housing are now required.

Animal Welfare↗

Calf response to caustic paste and hot-iron dehorning using sedation with and without local anesthetic.

Dairy producers require effective methods of reducing the pain associated with dehorning. Previous work has shown that analgesics can reduce pain associated with hot-iron dehorning, but these interventions may not be practical for some producers. Some producers favor the use of caustic paste (to cause a chemical burn as opposed to thermal burn), but little is known about how to treat pain caused by caustic burns. In 2 experiments, head shaking and head rubbing behaviors were used to evaluate pain following dehorning using caustic paste. In experiment 1, sedated calves were dehorned using a caustic paste, with or without a lidocaine local block. Calves treated with lidocaine showed no evidence of reduced pain response in the hours after the caustic paste was applied. In experiment 2, response to caustic paste dehorning with a sedative only was compared with hot-iron dehorning using a sedative and local anesthetic. In the first 4 h after dehorning, calves dehorned with a hot iron showed more head shaking than calves dehorned with caustic paste. These results indicate that dehorning with caustic paste combined with a sedative results in less pain to calves than dehorning with a hot iron combined with a sedative and a local anesthetic.

Adrenergic alpha-Agonists↗

Short communication: Effect of feed barrier design on the behavior of loose-housed lactating dairy cows.

The objective of this study was to evaluate the effects of 2 feed barrier systems on feeding and social behavior of dairy cows. Forty-eight lactating Holstein cows were subjected to each of 2 treatments in a cross-over design. The treatments were 2 types of feed-line barriers: 1) post-and-rail, and 2) headlock. Time-lapse video was used to quantify the feeding behavior and incidence of aggressive displacements of the cows at the feed bunk. Average daily feeding time did not differ when cows used the headlock barrier compared with the post-and-rail barrier. However, there were certain changes in feeding time during periods of peak feeding activity: cows that had lower feeding times relative to group mates when using the post-and-rail barrier showed more similar feeding times to group mates when using the headlock barrier. There were 21% fewer displacements at the feed bunk when cows accessed feed by the headlock barrier compared with the post-and-rail barrier. These results suggest that using a headlock barrier reduces aggression at the feed bunk and improves access to feed for socially subordinate cows during peak feeding periods.

Aggression↗

Freestall maintenance: effects on lying behavior of dairy cattle.

In a series of 3 experiments, we documented how sand-bedding depth and distribution changed within freestalls after new bedding was added and the effect of these changes on lying behavior. In experiment 1, we measured changes in bedding depth over a 10-d period at 43 points in 24 freestalls. Change in depth of sand was the greatest the day after new sand was added and decreased over time. Over time, the stall surface became concave, and the deepest part of the stall was at the center. Based on the results of experiment 1, we measured changes in lying behavior when groups of cows had access to freestalls with sand bedding that was 0, 3.5, 5.2, or 6.2 cm at the deepest point, below the curb, while other dimensions remained fixed. We found that daily lying time was 1.15 h shorter in stalls with the lowest levels of bedding compared with stalls filled with bedding. Indeed, for every 1-cm decrease in bedding, cows spent 11 min less time lying down during each 24-h period. In a third experiment, we imposed 4 treatments that reflected the variation in sand depth within stalls: 0, 6.2, 9.9, and 13.7 cm below the curb. Again, lying times reduced with decreasing bedding, such that cows using the stalls with the least amount of bedding (13.7 cm below curb) spent 2.33 h less time per day lying down than when housed with access to freestalls filled with sand (0 cm below curb).

Animals↗

Changes in feeding, drinking, and standing behavior of dairy cows during the transition period.

The aims of this study were to determine how measures of feeding, drinking, and standing behavior change over the period around calving, to derive objective meal and drinking bout criteria for transition cows, and to describe the within-cow consistency of these behavioral measures. Fifteen transition dairy cows were monitored from 10 d before until 10 d after calving. Meal criteria were calculated for each cow in both the pre- and postcalving periods and varied among cows from 5.3 to 105.2 min. There was a tendency for the average number of meals per day to be higher after calving than before calving, but time spent eating declined from 87 to 62 min/d from the precalving to the postcalving period. Time spent drinking before calving averaged 5.5 min/d and increased gradually after calving to an average of 6.8 min/d. Total daily standing times remained reasonably similar over the transition period but were highest at around calving (14.4 h) and lowest during the precalving period (12.3 h). On the day of calving, there was a dramatic increase in the number of standing bouts (21.8 bouts) compared with the pre- and postcalving averages of 11.7 and 13.1 bouts, respectively. In summary, changes in feeding behavior may help account for the well-documented changes in feed intake during transition. Documented changes in standing behavior suggest that cow comfort may be particularly important during the time around calving.

Animals↗

Influence of neck-rail placement on free-stall preference, use, and cleanliness.

Three experiments examined how the presence of a neck rail at different heights and locations influenced dairy cattle behavior and stall cleanliness. Experiment 1 compared 4 levels of neck-rail height (102, 114, and 127 cm and no neck rail; presented at 160 or 180 cm from the curb) in a preference test. Cows (n = 10) showed no consistent preference based on neck-rail height, regardless of the horizontal position of the neck rail. When cows were restricted to each treatment in turn, however, time spent standing fully (with all 4 hooves) in the stall was least in the stall with the lowest neck rail (mean, 22 min/24 h) and was greatest in the stall with no neck rail (mean, 83 min/24 h). A second experiment examined the effect of a neck rail placed at 3 distances from the curb (140, 175, and 233 cm) when height was held constant (131 cm; n = 12). Time spent standing fully in the stall was least when the neck rail was close to the curb (140 cm; mean, 11 min/24 h) and was greatest when the neck rail was furthest from the curb (233 cm; mean, 86 min/24 h). When the neck rail was far from the curb, the cows were more likely to soil the stall by defecating while standing fully in the stall. Experiment 3 compared soiling of the stall by 14 cows with and without a neck rail at a height of 124.5 cm. When the neck rail was removed, cows were more than twice as likely to soil the stall by defecating while standing fully in the stall compared with when the neck rail was present (1.3 vs. 0.5 defecations/24 h). Thus, restrictive neck-rail placement prevents cows from standing in stall, but helps keep stalls clean. Access to more comfortable flooring surfaces outside the stall may help mitigate the negative effects of restrictive neck rails.

Animals↗

Feeding behavior identifies dairy cows at risk for metritis.

Dairy cows experience a high incidence of disease and metabolic disorders in the weeks immediately following calving, but early and accurate diagnosis remains a challenge. Cows suffering from metritis, one common disease following calving, exhibit reduced milk yield and reproductive performance. However, afflicted cows show few overt signs of illness and frequently go unnoticed in the absence of veterinary examination. To determine whether changes in feeding behavior could be used to identify animals at risk for metritis, attendance at the feed alley was monitored continuously for 26 Holstein cows during the transition period, beginning 2 wk before and ending 3 wk after calving. Every 3 +/- 1 d, cows were examined for metritis based on rectal body temperature and condition of vaginal discharge. Over the 3 wk of observations after calving, 69% of cows showed some signs of metritis. These cows spent on average 22 min/d less time at the feed alley during the transition period than did non-metritic cows. For every 10-min decrease in average daily feeding time, cows were twice as likely to be diagnosed with metritis. A threshold of 75 min of average daily feeding time was 89% sensitive and 62% specific for detection of acute metritis. In conclusion, reduced time at the feeder can be used to identify dairy cows at risk for metritis. More research is required to determine how soon before calving at-risk cows can be identified and whether these behavioral differences can also be used in the early diagnosis of other diseases or metabolic disorders.

Animals↗

Hoof pathologies influence kinematic measures of dairy cow gait.

To explore how hoof pathologies affect the gait of dairy cattle, we studied gait profiles of cows with no visible injuries (n = 17), sole lesions (n = 14), and sole ulcers (n = 7). Video recordings of dairy cows were digitized using motion analysis software to calculate 6 stride variables for each hoof. Compared with cows with sole ulcers, healthy cows walked faster (1.11 +/- 0.03 vs. 0.90 +/- 0.05 m/s, mean +/- SEM), had shorter stride durations (1.26 +/- 0.03 vs. 1.48 +/- 0.05 s), and longer strides (139.5 +/- 2.1 vs. 130.0 +/- 3.2 cm). Percentage of triple support in the gait cycle (time when cattle were supported by 3 legs) more than doubled for cows with sole ulcers compared with healthy cows (42 vs. 18%). Gait differences were likely due to cows reducing the load on an affected leg. Few differences were detected between healthy cows and those with sole lesions, perhaps because of variation in number, severity, and location of injuries. Kinematic gait analysis is a promising approach in understanding how hoof pathologies affect dairy cow gait.

Animals↗

Improving stall design: use of 3-D kinematics to measure space use by dairy cows when lying down.

Uncomfortable stalls reduce the time cows spend resting and are a risk factor for lameness. Although recommendations for stall dimensions are available, little research supports these recommendations. In this study, kinematic techniques were used to provide the first accurate measures of space used by Holstein dairy cows during lying-down movements in an open space and in a free stall. Cows (n=5, mean parity 3.2 +/- 1.0, body weight=607 +/- 21 kg) used up to 300 cm of longitudinal space when lying down, more than is typically provided by current industry recommendations for stall length. Cows used up to 109 cm of lateral space (180% of hip width), an estimate that is within current recommendations for stall width. During the lying-down movement, maximal lateral displacements at the hip occurred in two vertical zones: one between 95 and 135 cm, and the second less than 50 cm above the lying surface. Maximal longitudinal displacements of the nose were clustered 10 to 30 cm above the surface. These heights should be taken into account when designing and positioning stall partitions. Maximum instantaneous velocity of body markers approached 220 cm/s, indicating that cows can contact inappropriately placed stall partitions and the lying surface with considerable force. Kinematic techniques are promising ways of assessing spatial requirements of cattle to improve stall design. Further work is required to assess a wider range of cow sizes and stall configurations.

Animals↗

Free-stall dimensions: effects on preference and stall usage.

In 2 experiments, free-stall dimensions were examined to determine how they affected stall preference, usage, cleanliness, and milk production in Holstein dairy cattle. In experiment 1, stall width (112 or 132 cm) and stall length (229 and 274 cm from curb to wall) were compared in a 2 x 2 factorial arrangement of stall treatments using 15 individually housed, non-lactating animals. Cows showed no clear preference for stall size as measured by lying time. When animals had no choice between stalls, average lying time was higher in the wide stalls than in the narrow stalls (10.8 vs. 9.6+/-0.3 h/24 h). Both length and width affected time spent standing with only the front hooves in the stall; total stall area is best explained by the variation associated with this behavior. In experiment 2, 27 lactating dairy cattle were alternately housed with access to stalls of 106, 116, or 126 cm in width using a cross-over design with exposure to each treatment lasting 3 wk. Animals spent an additional 42 min/24 h lying in stalls measuring 126 cm in width compared with stalls with only 106 cm between partitions. Free-stall width influenced the time spent standing with the front 2 hooves in the stall; animals averaged 58 min/24 h in the widest stalls and 85 min/24 h in the narrowest stalls. The amount of time spent standing with all 4 hooves in the stall tended to be longer in wider stalls, and these stalls were also most likely to become soiled with feces. Stall width did not affect the number of lying events or milk production. In conclusion, animals spent more time lying down, and less time was spent standing with only the front hooves in larger stalls, but larger stalls were also more likely to become soiled.

Animals↗

Effect of feeding space on the inter-cow distance, aggression, and feeding behavior of free-stall housed lactating dairy cows.

The objectives of this study were to determine whether doubling the amount of feeding space from 0.5 to 1.0 m per cow leads to increased spacing between cows at the feeder, fewer aggressive social interactions among cows, and ultimately increased feeding activity. Twenty-four lactating Holstein cows were provided with 0.5 and 1.0 m of feeding space per cow in a 2 x 2 crossover design replicated over time. Time-lapse video was used to quantify the inter-cow distance and incidence of aggressive displacements at the feed alley. An electronic feed alley monitoring system was used to monitor the feeding behavior of the cows. When animals had access to 1.0 m per cow, there was at least 60% more space between animals and 57% fewer aggressive interactions while feeding than with access to 0.5 m of feeding space. These changes in spacing and aggressive behavior in turn allowed cows to increase feeding activity throughout the day, especially during the 90 min after providing fresh feed (an increase of 24%). This increase in feeding activity was particularly evident for subordinate cows. These results indicate that increasing space allowance at the feeder increases feeding activity and reduces competition among lactating dairy cows.

Aggression↗